PLSC 210: Lecture 4

GROWTH AND METABOLISM

I.  TERMS
        1. Growth- Irreversible increase in size

2. Development-
Morphogenesis - Morphological and anatomical development
Differentiation - Physiological and biochemical specialization of plant tissues
3. Metabolism- Synthesis an degradation of organic compounds
Anabolism - Synthesis
Catabolism - Degradation
II. MAJOR CHEMICAL PROCESSES OF PLANTS
1. Photosynthesis
                                    Chloroplast
12H2O + 6CO2 + Light Clorophyll >C6H12O6 + 6O2 + 6H2O
(water)+ (Carbon Dioxide)+(energy)------->  (Carboydrate)+(oxygen)+(water)
2. Metabolism
                              Cytoplasm
C6H12O6 + Mineral  Enzymes > Various organic compounds
(Carbohydrate) + (fertilizer)----> (Protiens, fats, starch, hormones, vit, etc.)
3. Respiration
                                        Mitochondria
Organic compounds + O2 Enzymes > CO2 + H2O + Energy + Mineral
(Substates)             (oxygen)                                          (ATP)     (inorganic)
energy source

The energy released fron respiration is used for growth and development

III. PHOTOSYNTHESIS

          The process of producing carbohyrates from CO2 and water by using light energy in the presence of chlorophyll.  Takes place in the chloroplast.
        1. Light Phase of Photosynthesis

         Photolysis-cleavage of water intao hydrogen and oxygen by light (Hill Reaction
         
     

         Photophosphorylation- Conversion of ADP to ATP by light energy
                

     
     

          Sum:   conversion of light energy to chemical energy
          

     2. Dark Phase of Photosynthesis

         The Calvin Cycle- a series of enzymaticaly mediated reactions in which CO2 is reduced to 3-phosphoglyceraldehyde (3PGA) and the CO2 receptor (ribulose biphosphate, RUBP) is generated
     

                      
     
     

        The Net Gain
 
                                     Chloroplast
12H2O + 6CO2 + Light Clorophyll >C6H12O6 + 6O2 + 6H2O
        3. Two Different CO2 Pathways
               The C3 Pathway
                - Occurs in C3 plants (many dicots like soybean, tomato, apple)
                - The first product of CO2 fixation is C3 acids
                - Only the Calvin Cycle operates
                - Photorespiration exists
                                

               The C4 Pathway
                - Occurs in C4 plants (tropical grasses, corn, sugarcane, some dicots like amaranth, atriplex)
                - The first product of CO2 fixation is C4 acids
                - Both the Calvin Cycle and C pathway operate
                - Lacks photorespiration

                     
 

                     
 

        4. Photorespiration
                The process of respiration that consumes oxygen and releases CO2 in the presence of light
            -Consumes the reducing power to reduce O2 to CO2
                 -Does not produce ATP
                 -Reduces photosynthetic efficiency
                 -Occurs in C3 plants

             

    5. Carbon Dioxide Compensation Point
         A steady state of CO2 concentration in the air at which CO2 taken up by plants via photosynthesis is the same as the CO2 given off via respiration
         - At COcompensation point, no growth occurs
         - Below compensation point, plants will degrade
         - C3 plants have higher CO2 compensation points than the C4 plants
          COCompensation points
            Soybean (C3 plant)-----50ppm at 25° C
            Corn (C4 plant)---------10ppm at 25°C
                    Ambient CO2------300ppm (0.03%)
        - Same principles apple to light compensation points
           Net Photosynthesis (PHS) = Gross PHS - Respiration
IV.  NUTRIENT ABSORPTION AND TRANSLOCATION
 

        1. Plant Nutrients

        -16 elements
          Macro nutrients:  N, P, K, Ca, Mg, S, C, O, H
          Micro nutrients:   B, Cl, Cu, Fe, Mn, Mo, Zn

    2. Ability to Manufacture Food
        - Most green plants are autotrophic
    Autotrophic- Capable of manufacturing its own food from minerals
    Heterotrophic- Incapable of manufacturing its own food, so the plant must depend on other sources for organic matter
                          (immature embryo, dodder, human)

    3. Nutrient and Water Movement
        -Diffusion- Movement of molecules (substance) from a region of high concentration to the region of low concentration
        -Osmosis- Diffusion of water through differentially permeable membrane
        -Translocation- Movement of inorganic and organic solutes from one part of plant to another
                              - Water conduction and mineral movement via xylem
                              - Carbohydrate translocation through phloem
        -Transpiration- loss of water vapor from the leaf via Stomata
        -Evaporation- liquid water turns into water vapor
        -Evapotraspiration- Evaporation and transpiration

V.  RESPIRATION

        1. The Reverse of Photosynthesis
         The process of releasing energy, carbon dioxide and water by oxidation of organic compounds

                C6H12O6 + 6O2         >6H2O + 6CO2 + Energy

    2. Chemical Processes
        Glycolysis- conversion of C6 sugars to CO2 and Pyruvic acid
         Citric acid cycle (Kreb Cycle)- oxydation of Pyruvic acid to H+ , e-, and CO2 in mitochondria
    3. The Q10
         -The rate of respiration doubles when temperature rises 10° C (18° F)
         - Respiration can be reduced by lowering 02 and increasing CO2 concentration

         Application
         - CO2 storage of apples, pears
         - Hypobaric storage of fruits and flowers
                 lower atmospheric pressure (vacuum)
                 lower temperature
     

VI.  METABOLISM

            Synthesis and degradation of organic compounds

            Anabolism- synthesis process

            Catabolism- Degradation or breakdown process (occurs in cytoplasm)

VII.  PLANT CONSTITUENTS

       1. Carbohydrates
             -Monosaccharides- simple carbohydrated (C5 Pentase, C6 hexose)
             -Disaccharides- C12 {i.e. maltose (glu-glu), sucrose (glu-fru)}
             -Olygosaccharides- 1-10 monosaccarides linked together
             -Polysaccharides- starch (poly glu), cellulose (poly glu), insulin hemicellulose

       2. Lipids
             -phospholipids, fats, waxes

        3. Proteins
             -structural, soluble (enzymes)
        4. Other Organic Compounds
           Aromatic compounds (vanillin, flavonoids, etc.)
           Terpenoids and steroids (brassinolids, GA metabolites)
           Non-protein nitrogen compounds (DNA, RNA, bases)
           Vitamins (Vitamin C, thiamin B, provitamin A, etc.)
           Others
           Pharmaceuticals [taxol (pacific yew), Echinacin, ginsengoside]