Agonoscelis Spinola,
1837
Agonoscelis versicoloratus
Isalo, Madagascar, December 2,
2013
Photo by Jean Brodeur
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Agonoscelis
Spinola, 1837: 327-329.
Nevroscia
Amyot & Serville, 1843: XXIII, 109. (syn. by Dallas, 1851)
Type Species:
Agonoscelis:
Agonoscelis indica Spinola, 1837 (= Cimex nubilis Fabricius, 1775),
by monotypy. Nevroscia: Pentatoma grata Palisot de
Beauvois, 1811 (= Cimex nubilis Fabricius, 1775), by subsequent
designation (Kirkaldy, 1903).
Tribal Placement:
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Included Species:
antennata Distant, 1911
arabica Linnavuori, 1975
bicolor Distant, 1879
campbelli Distant, 1918
cognata Horváth, 1904
erosa erosa (Westwood, 1837)
erosa atropurpurea Schumacher, 1913
femoralis Walker, 1868
haraldi Bergroth, 1908
karachiensis Ahmad, Siddiqui, & Kamaluddin, 2001
longirostris Harold, 1879
marshalli Schouteden, 1912
neavei Distant, 1910
nubilis (Fabricius, 1775)
odendaali Distant, 1910
okuana Linnavuori, 1982
puberula Stål, 1853
rutila (Fabricius, 1775) [image]
sansibarica Harold, 1879
tamilnadensis Mathew, 1977
vanderysti Schouteden, 1916
venosa (Thunberg, 1783)
versicoloratus fallaciosa Linnavuori, 1975
versicoloratus gambiensis (Westwood, 1837)
versicoloratus tibialis Horváth, 1904
versicoloratus versicoloratus (Turton, 1802)Comments:
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Key to species of Agonoscelis
(modified from Jensen-Haarup, 1920)
1
Body oblong-oval; abdominal venter (except spiracles) without black spots;
rostrum reaching beyond apex of abdomen (Angola to Sudan) |
longirostris Harold |
-
Body more or less broadly oval; abdominal venter with black spots arranged
in rows |
2 |
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2(1)
Body reddish, spotted with black; rostrum not reaching beyond middle of
second abdominal segment (Madagascar) |
bicolor Distant |
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Surface of body fuscous or rufous-testaceous, or testaceous spotted with
dark areas; rostrum reaching to or near middle of abdominal venter, or
even to apex of venter |
3 |
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3(2)
Corium equally punctate, without scattered, distinct, pale callosities |
4 |
-
Corium unequally punctate, with scattered, pale callosities |
7 |
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4(3)
Body narrowly oval; superior surfaces of tibiae black at base |
5 |
-
Body broadly oval; tibiae entirely pale |
6 |
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5(4)
Rostrum reaching to apex of abdominal venter; head longer than pronotum;
second and third segments of antennae subequal in length; dorsum of
abdomen reddish testaceous, with base and apex black; 10.5-12 mm in length
(Kenya, Tanzania) |
cognata Horváth |
-
Rostrum not reaching to apex of abdomen; head and pronotum of equal
length; second segment of antennae distinctly shorter than third; dorsum
of abdomen entirely rufous testaceous; 10-11 mm in length (Ethiopia to
South Africa) |
venosa (Thunberg) |
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6(4)
Dorsum of abdomen rufous testaceous; apex of scutellum reflexed; head with
eyes longer than broad; second segment of antennae distinctly shorter than
third; tarsi pale; 9.5 mm in length (Eritrea to Tanzania) |
sansibarica Harold |
-
Dorsum of abdomen black; apex of scutellum not reflexed; head not longer
than broad; second segment of antennae not shorter than third; tarsi more
or less blackish or fuscous; 8.5-9 mm in length (Eritrea to South Africa) |
puberula Stål |
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7(3)
Membrane (with exception of the usual, dark spot at inner angle) hyaline
and with veins pale; dorsum of abdomen black; 14-14.5 mm in length (Sierra
Leone to South Africa) |
erosa (Westwood) |
-
Membrane sordid hyaline, with fuscous or brownish veins; dorsum of abdomen
entirely or for the most part reddish |
8 |
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8(7)
Tibiae entirely black; 11.5-14 mm |
versicoloratus gambiensis (Westwood) |
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Tibieae at least partly pale |
9 |
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9(8)
Dorsum of abdomen entirely reddish; second and third segments of antennae
slender; equal in length; head long and strongly tapering anteriorly; head
and pronotum strongly pilose; legs entirely pale; 11-12 mm in length
(Nigeria to Ethiopia) |
haroldi Bergroth |
-
Dorsum of abdomen reddish, with base and apex black; second and third
segments of antennae thicker, second segment distinctly shorter than
third; head shorter, not strongly tapering anteriorly; head and pronotum
not strongly pilose; tarsi and often part of tibiae fuscous or blackish;
11-14 mm in length (Eritrea to South Africa) |
versicoloratus (Turton) |
Key to West African species of Agonoscelis
(modified from Linnavuori, 1982)
1
Dorsum of abdomen totally black; last tergite of
male with long median process directed caudad
(Sierra Leone to South Africa) |
erosa (Westwood) |
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Dorsum of abdomen not totally black; last tergite of male unarmed |
2 |
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2(1)
Hair covering of head, pronotum and scutellum very long, erect; hemelytra
also with distinct longish semidecumbent hairs; legs usually almost
totally pale; dorsum of abdomen generally uniformly red; paramere with
subapical tooth on outer margin |
haroldi Bergroth |
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Hair covering of head, pronotum and scutellum moderately long, erect, that
of hemelytra inconspicuous; dorsum of abdomen bicolored; legs bicolored;
apex of paramere T-shaped |
3 |
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3(2)
Length 11-14 mm; head about 1.2 times as long as broad, lateral margins
nearly always distinctly insinuated in front of eyes; scutellum 1.3-1.4
times as long as broad, apical part usually narrow ligulate; lateral lobes
of pygophore in lateral view narrow, concealed by bluntly produced
elevated lateral margin of pygophore basally; paramere with T-shaped apex
(both apical angles sharp) |
versicoloratus (Turton) |
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Length 10-12.5 mm; head appearing short, 1.0-1.12 times as long as broad,
lateral margins not insinuated in front of eyes; scutellum broader,
tapering more gradually apicad; lateral lobes of pygophore in lateral view
broadly visible; lower part of lateral margin of pygophore forming a
ligulate process; paramere with only one of the apical angles sharp |
okuana Linnavuori |
Other Links:
Stink Bugs of Australia
References:
Jensen-Haarup, A. C. 1920.
Hemipterological notes and descriptions I. Entomologiske Meddelelsen 13:209-224.
Linnavuori, R. E. 1982. Pentatomidae and
Acanthosomatidae (Heteroptera) of Nigeria and the Ivory Coast, with remarks on
species of the adjacent countries in West and Central Africa. Acta Zoologica
Fennica, no. 163: 176 pp.
David A. Rider
Professor of Entomology
North Dakota State University
202 Hultz Hall
Fargo, ND 58105
E-Mail:
David.Rider@ndsu.edu
updated:
26 Feb 2015
Published by the
Department of
Entomology
Prospective students may schedule a visit by calling 1-800-488-NDSU.
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