Tolumnia Stål, 1867
Tolumnia Stål,
1867: 515.
Mormoschema
Breddin, 1909: 269-270. (syn. by Distant, 1918)
Type Species:
Tolumnia:
Pentatoma trinotata Westwood, 1837, by subsequent designation (Distant,
1902). Mormoschema: Tolumnia immaculata Distant, 1900, by
monotypy.
Included Species:
antennata Distant, 1902
basalis (Dallas, 1851)
elongata Hasan & Ahmad, 1988
gutta (Dallas, 1851)
horni Breddin, 1909
immaculata Distant, 1900
latipes (Dallas, 1851)
longirostris (Dallas, 1851)
malayensis Hasan & Ahmad, 1988
maxima Distant, 1902
papulifera Bergroth, 1922
southwoodi Hasan & Ahmad, 1988
trinotata (Westwood, 1837)
Comments:
After studying many specimens belonging to the genus
Tolumnia, I now believe that I can identify most of the common Tolumnia
species. I have examined specimens of the following species:
1) antennata Distant, 1902
(type locality = India)
2) basalis (Dallas, 1851)
(China)
3) gutta (Dallas, 1851)
(China)
4) latipes (Dallas, 1851) (Tenasserim)
5) longirostris (Dallas, 1851)
(Philippines)
6) trinotata (Westwood, 1837)
(Java)
Species I have not examined:
1) elongata Hasan & Ahmad,
1988 (Malaysia)
2) horni Breddin, 1909 (Sri
Lanka)
3) immaculata Distant, 1900
(India, Sri Lanka)
4) malayensis Hasan & Ahmad,
1988 (Sumatra)
5) maxima Distant, 1902 (Tenasserim)
6) papulifera Bergroth, 1922
(Sarawak)
7) southwoodi Hasan & Ahmad,
1988 (Malaya)
Hasan &
Ahmad’s species were based primarily on differences in the spermathecal
bulb (having one finger-like projection versus two or more; this is a character
that can vary within a single species). Breddin indicated that immaculata
may have a shorter ostiolar peritreme (I need to check this). Distant described
maxima as being larger, and somewhat similar to Halyomorpha in
size and appearance. I have not checked the descriptions of horni or
papulifera yet.
Key to the species of Tolumnia I am familiar
with
1
Rostrum elongate, reaching well onto the fourth or fifth abdominal
sternite; relatively large medial pale spot on anterior disk of pronotum
(Philippines) |
longirostris (Dallas) |
-
Rostrum much shorter, reaching to or at most slightly beyond posterior
margin of hind coxae; anterior disk of pronotum lacking medial pale spot,
at most irregularly and only slightly pale in this area |
2 |
|
|
2(1) All
antennal segments black, although basal areas of some segments may be pale
(India) |
antennata
Distant |
-
At least segments two and three of antennae pale, segment one usually
pale, but may be darkish, segments four and five usually blackish, but
each with pale basal area |
3 |
|
|
3(2)
Tylus usually nearly impunctate, paler in color than juga; apices of
anterior tibiae usually distinctly dilated; relatively large pale spot in
each basal angle of scutellum; usually with relatively large black spots
or bands on legs, especially on joint between femur and tibia, and on apex
of each tibia (southeast Asia, Malaysia, western Indonesia) |
latipes (Dallas) |
–
Tylus usually with some punctures, nearly concolorous with juga; apices of
anterior tibiae usually not dilated, if dilated, then only slightly so;
basal angles of scutellum with or without pale spots; legs often speckled
with brown or black, but usually lacking well defined black spots or bands
as described above |
4 |
|
|
4(3)
Basal angles of scutellum with relatively large pale spots (Java) |
trinotata (Westwood) |
–
Basal angles of scutellum either lacking pale spots, spots very tiny, or
pale area extensive, forming irregular band across base of scutellum |
5 |
|
|
5(4)
Antennal segment five black on apical half, pale on basal half; dark bands
on connexiva relatively broad, each connexival segment divided into thirds
- black, pale, black (China, Taiwan) |
gutta
(Dallas) |
–
Antennal segment five black with pale area limited to basal one fourth or
less; dark bands on connexiva relatively narrow, pale area taking up at
least half of each connexival segment (China, India, Thailand,
Philippines) |
basalis (Dallas) |
David A. Rider
Associate professor of Entomology
NDSU
202 Hultz Hall
Fargo, ND 58105
E-Mail: David.Rider@ndsu.nodak.edu
Published by the Department of
Entomology
Prospective students may schedule a visit by calling 1-800-488-NDSU.
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